
Publications
Miyashita, T., Janvier, P., Tietjen, K., Berenguer, F., Schöder, S., Marone, F., Gueriau, P., Coates, M. 2025. Novel assembly of a head–trunk interface in the sister group of jawed vertebrates. Nature 645:686-691. doi:10.1038/s41586-025-09329-9
Montheil, L., Licht, A., İbilioğlu, D., Botté, P., Ocakoğlu, F., Demory, F., Ruffet, G., Guihou A., Kaya, M., Raynaud B., Akkiraz, M.S., Deschamps, P., Métais, G., Coster, P., and Beard, K.C. 2025. Updating the timeline of faunal endemism in Balkanatolia, the biogeographic province connecting Europe, Asia and Africa. Journal of Asian Earth Sciences 290: 106661. doi:10.1016/j.jseaes.2025.10661
Crowell, J.W., Beard, K.C., and Chester, S.G.B. 2025. Micro-computed tomography unveils anatomy of the oldest known plesiadapiform cranium. Journal of Human Evolution 201: 103655. doi:10.1016/j.jhevol.2025.103655
Mulcahy, K.D., Constenius, K. N., and Beard, K.C. 2025. Northernmost record of Dinocerata (Mammalia: Eutheria) in North America from the Middle Eocene Kishenehn Formation of Montana. Annals of Carnegie Museum 90: 3. doi:10.2992/007.090.0305
Miller, K., and Beard, K.C. 2025. Biogeographic and Biostratigraphic Implications of a New Species of Swaindelphys (Mammalia, Metatheria) from the Paleocene (Tiffanian) Black Peaks Formation, Big Bend National Park, Texas. Journal of Vertebrate Paleontology 44: 6. doi:10.1080/02724634.2025.2500501
Jones, M.F., Beard, K.C., and Simmons, N.B. 2024. Phylogeny and systematics of early Paleogene bats. Journal of Mammalian Evolution 31: 18. doi:10.1007/s10914-024-09705-8
Magallanes, I., Beard, K.C., Martin, T., and Luo, Z.-X. 2024. A new dryolestid fossil from the Late Jurassic illuminates molar root structure of dryolestids. Journal of Vertebrate Paleontology 43: e2322740. doi:10.1080/02724634.2024.2322740
Métais, G., Coster, P., Kaya, M., Licht, A., Miller, K., Ocakoğlu, F., Rust, K., and Beard, K.C. 2024. Rapid colonization and diversification of a large-bodied mammalian herbivore clade in an insular context: New embrithopods from the Eocene of Balkanatolia. Journal of Mammalian Evolution 31: 15. doi:10.1007/s10914-024-09711-w
Beard, K.C., and Métais, G. 2024. Oldest record of Apatemyidae (Mammalia, Apatotheria) from Spain and the taxonomic status of Spanish paromomyids (Mammalia, Primatomorpha). Journal of Vertebrate Paleontology 39: e1730389. doi:10.1080/02724634.2023.2288651
Anemone, R.L., Jones, M.F., Van Regenmorter, J., Beard, K.C. 2024. A latest Paleocene mammal fauna from the Great Divide Basin in southern Wyoming and a revised biozonation of the Clarkforkian land mammal age. Journal of Vertebrate Paleontology 44: 3. doi:10.1080/02724634.2024.2424139
Métais, G., Coster, P.M.C., Licht, A., Ocakoğlu, F., and Beard, K.C. 2023. Additions to the late Eocene Süngülü mammal fauna in easternmost Anatolia and the Eocene-Oligocene transition at the periphery of Balkanatolia. Comptes Rendus Palevol 22: 711-727. doi:10.5852/cr-palevol2023v22a35
Rust, K., Ni, X., Tietjen, K., and Beard, K.C. 2023. Phylogeny and paleobiogeography of the enigmatic North American primate Ekgmowechashala illuminated by new fossils from Nebraska (USA) and Guangxi Zhuang Autonomous Region (China). Journal of Human Evolution 185: 103452. doi:10.1016/j.jhevol.2023.103452
Jones, M.F., and Beard, K.C. 2023. Nyctitheriidae (Mammalia, ?Eulipotyphla) from the late Paleocene of Big Multi Quarry, southern Wyoming, and a revision of the subfamily Placentidentinae. Annals of Carnegie Museum 88: 115-159. doi:10.2992/007.088.0202
Beard, K.C., Coster, P.M.C., Ocakoğlu, F., Licht, A., and Métais, G. 2023. Dental anatomy, phylogenetic relationships and paleoecology of Orhaniyeia nauta (Metatheria, Anatoliadelphyidae), a Gondwanan component of the insular Eocene mammal fauna of Balkanatolia (north-central Turkey). Journal of Mammalian Evolution 30: 859-872. doi:10.1007/s10914-023-09680-6
Caron, A., Venkataraman, V., Tietjen, K., and Coates, M. 2023. A fish for Phoebe: a new actinopterygian from the Upper Carboniferous Coal Measures of Saddleworth, Greater Manchester, UK, and a revision of Kansasiella eatoni. Zoological Journal of the Linnean Society 198(4):1-25. doi:10.1093/zoolinnean/zlad011
Miller, K., Tietjen, K., and Beard, K.C. 2023. Basal Primatomorpha colonized Ellesmere Island (Arctic Canada) during the hyperthermal conditions of the early Eocene climatic optimum. PLoS ONE 18: e0280114. doi:10.1371/journal.pone.0280114
Campbell, C.F., Mueller, M.A., Taylor, M.H., Ocakoğlu, F., Möller, A., Métais, G., Coster, P.M.C., Beard, K.C., and Licht, A. 2023. The geodynamic implications of passive margin subduction in northwest Turkey. Geochemistry, Geophysics, Geosystems 24: e2022GC010481. doi:10.1029/2022GC010481
Mueller, M.A., Licht, A., Campbell, C., Ocakoğlu, F., Akşit, G.G., Métais, G., Coster, P.M.C., Beard, K.C., and Taylor, M.H. 2022. Sedimentary provenance from the evolving forearc-to-foreland central Sakarya Basin, western Anatolia reveals multi-phase intercontinental collision. Geochemistry, Geophysics, Geosystems 23: e2021GC010232. doi:10.1029/2021GC010232
Miyashita, T., Gess, R.W., Tietjen, K., and Coates, M.I. 2021. Non-ammocoete larvae of Palaeozoic stem lampreys. Nature 591: 408–412. doi:10.1038/s41586-021-03305-9
Licht, A., Métais, G., Coster, P., Ibilioğlu, D., Ocakoğlu, F., Westerweel, J., Mueller, M., Campbell, C., Mattingly, S., Wood, M.C., and Beard, K.C. 2022. Balkanatolia: The insular mammalian biogeographic province that partly paved the way to the Grande Coupure. Earth-Science Reviews 226: 103929. doi:10.1016/j.earscirev.2022.103929
Jones, M.F., Li, Q., Ni, X., and Beard, K.C. 2021. The earliest Asian bats (Mammalia: Chiroptera) address major gaps in bat evolution. Biology Letters 17: 20210185. doi:10.1098/rsbl.2021.0185
Boyer, D.M., Schaeffer, L.M., and Beard, K.C. 2021. New dentaries of Chiromyoides (Primatomorpha, Plesiadapidae) and a reassessment of the “mammalian woodpecker” ecological niche. Geobios 66-67: 77-102. doi:10.1016/j.geobios.2021.03.002
Mattingly, S.G., Beard, K.C., Salem, M.J., Chaimanee, Y., and Jaeger, J.-J. 2021. A new parapithecine (Primates: Anthropoidea) from the early Oligocene of Libya supports parallel evolution of large body size among parapithecids. Journal of Human Evolution 153: 102957. doi:10.1016/j.jhevol.2021.102957
Mattingly, S.G., Beard, K.C., Coster, P.M.C., Salem, M.J., Chaimanee, Y., and Jaeger, J.-J. 2020. A new carnivoraform from the early Oligocene of Libya: Oldest known record of Carnivoramorpha in Africa. Journal of African Earth Sciences 172: 103994. doi:10.1016/j.jafrearsci.2020.103994
Beard, K.C., Métais, G., Ocakoğlu, F., and Licht, A. 2021. An omomyid primate from the Pontide microcontinent of north-central Anatolia: Implications for sweepstakes dispersal of terrestrial mammals during the Eocene. Geobios 66-67: 143-152. doi:10.1016/j.geobios.2020.06.008
Beard, K.C., Jones, M.F., Thurber, N.A., and Sanisidro, O. 2020. Systematics and paleobiology of Chiromyoides (Mammalia, Plesiadapidae) from the Upper Paleocene of western North America and western Europe. Journal of Vertebrate Paleontology 39: e1730389. doi:10.1080/02724634.2019.1730389
Mueller, M.A., Licht, A., Campbell, C., Ocakoğlu, F., Taylor, M.H., Burch, L., Ugrai, T., Kaya, M., Kurtoğlu, B., Coster, P.M.C., Métais, G., and Beard, K.C. 2019. Collision chronology along the İzmir-Ankara-Erzincan suture zone: insights from the Sarıcakaya Basin, Western Anatolia. Tectonics 38: 3652-3674. doi:10.1029/2019TC005683
Coster, P.M., Soe, A.N., Beard, K.C., Chaimanee, Y., Sein, C., Lazzari, V., and Jaeger, J.-J. 2019. Astragalus of Pondaungimys (Rodentia, Anomaluroidea) from the late middle Eocene Pondaung Formation, central Myanmar. Journal of Vertebrate Paleontology 36: e1552156. doi:10.1080/02724634.2018.1552156
Jones, M.F., Coster, P.M.C., Licht, A., Métais, G., Ocakoğlu, F., Taylor, M.H., and Beard, K.C. 2019. A stem bat (Chiroptera: Palaeochiropterygidae) from the late middle Eocene of northern Anatolia: implications for the dispersal and palaeobiology of early bats. Palaeobiodiversity and Palaeoenvironments 99: 261-269. doi:10.1007/s12549-018-0338-z
Coates, M.I., Tietjen, K., Olsen, A.M., and Finarelli, J.A. 2019. High-performance suction feeding in an early elasmobranch. Science Advances 5(9): 1-8. doi:10.1126/sciadv.aax2742
Coates, M.I., and Tietjen, K. 2018. ‘This strange little palaeoniscid’: a new early actinopterygian genus, and commentary on pectoral fin conditions and function. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 109(1-2): 15-31 doi:10.1017/S1755691018000403
Métais, G., Coster, P.M., Kappelman, J.R., Licht, A., Ocakoğlu, F., Taylor, M.H., and Beard, K.C. 2018. Eocene metatherians from Anatolia illuminate the assembly of an island fauna during Deep Time. PLoS ONE 13: e0206181. doi:10.1371/journal.pone.0206181
Rossie, J.B., Smith, T.D., Beard, K.C., Godinot, M., and Rowe, T.B. 2018. Nasolacrimal anatomy and haplorhine origins. Journal of Human Evolution 114: 176-183. doi:10.1016/j.jhevol.2017.11.004
Coates, M., Finarelli, J., Sansom, I., Andreev, P., Criswell, K., Tietjen, K., La Riviere, P. 2018. An early chondrichthyan and the evolutionary assembly of a shark body plan. Proceedings of the Royal Society B: Biological Sciences 285: 20172418. doi:10.1098/rspb.2017.2418
Gebo, D.L., Dagosto, M., Ni, X., and Beard, K.C. 2017. Phalangeal morphology of Shanghuang fossil primates. Journal of Human Evolution 113: 38-82. doi:10.1016/j.jhevol.2017.08.001
Mattingly, S.G., Sanisidro, O., and Beard, K.C. 2017. A new species of Carpolestes (Mammalia, Plesiadapoidea) from the late Paleocene of southern Wyoming: assessing changes in size and shape during the evolution of a key anatomical feature. Historical Biology 30: 1031-1042. doi:10.1080/08912963.2017.1328509
Licht, A., Coster, P., Ocakoğlu, F., Campbell, C., Métais, G., Mulch, A., Taylor, M., Kappelman, J., and Beard, K.C. 2017. Tectono-stratigraphy of the Orhaniye Basin, Turkey: implications for collision chronology and Paleogene biogeography of central Anatolia. Journal of Asian Earth Sciences 143: 45-58. doi:10.1016/j.jseaes.2017.03.033
Métais, G., Erdal, O., Erturaç, K., and Beard, K.C. 2017. Tarsal morphology of the pleuraspidotheriid mammal Hilalia from the middle Eocene of Turkey. Acta Palaeontologica Polonica 62: 173-179. doi:10.4202/app.00314.2016
Coates, M., Gess, R., Finarelli, J., Criswell, K., and Tietjen, K. 2017. A symmoriiform chondrichthyan braincase and the origin of chimaeroid fishes. Nature 541: 208-211. doi:10.1038/nature20806
Gebo, D.L., Dagosto, M., Beard, K.C., and Ni, X. 2017. Cuboid morphology of a basal anthropoid from the Eocene of China. Journal of Human Evolution 102: 72-74. doi:10.1016/j.jhevol.2016.10.003
Coates, M.I., and Tietjen, K., 2017. The neurocranium of the Lower Carboniferous shark Tristychius arcuatus (Agassiz, 1837). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 108(1): 19-35. doi:10.1017/S1755691018000130
Beard, K.C. 2016. Out of Asia: anthropoid origins and the colonization of Africa. Annual Review of Anthropology 45: 199-213. doi:10.1146/annurev-anthro-102215-100019
Beard, K.C., Ni, X., Wang, Y.-Q., Meng, J., and Gebo, D.L. 2016. Dentition of Subengius mengi (Mammalia: Plesiadapoidea) and a reassessment of the phylogenetic relationships of Asian Carpolestidae. Vertebrata PalAsiatica 54: 181-211.
Ni, X., Li, Q., Li, L., and Beard, K.C. 2016. Oligocene primates from China reveal divergence between African and Asian primate evolution. Science 352: 673-677. doi:10.1126/science.aaf2107
Beard, K.C., Coster, P.M.C., Salem, M.J., Chaimanee, Y., and Jaeger, J.-J. 2016. A new species of Apidium (Anthropoidea, Parapithecidae) from the Sirt Basin, central Libya: First record of Oligocene primates from Libya. Journal of Human Evolution 90: 29-37. doi:10.1016/j.jhevol.2015.08.010
Beard, K.C., and Coster, P.M. 2016. Upper molar morphology of the early Oligocene Egyptian anthropoid Qatrania wingi. American Journal of Physical Anthropology 159: 714-721. doi:10.1002/ajpa.22913
Coster, P.M.C., Beard, K.C., Salem, M.J., Chaimanee, Y., and Jaeger, J.-J. 2015. New fossils from the Paleogene of central Libya illuminate the evolutionary history of endemic African anomaluroid rodents. Frontiers in Earth Science 3: 56. doi:10.3389/feart.2015.00056
Gebo, D.L., Beard, K.C., Ni, X., and Dagosto, M. 2015. Distal phalanges of Eosimias and Hoanghonius. Journal of Human Evolution 86: 92-98. doi:10.1016/j.jhevol.2015.05.014
Coster, P.M.C., Beard, K.C., Salem, M.J., Chaimanee, Y., Brunet, M., and Jaeger, J.-J. 2015. A new early Oligocene mammal fauna from the Sirt Basin, central Libya: biostratigraphic and paleobiogeographic implications. Journal of African Earth Sciences 104: 43-55. doi:10.1016/j.jafrearsci.2015.01.006